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Breeder's Equation Calculator (Response to Selection)

Predict genetic gain with the breeder's equation R = h²S — the response per generation from heritability and the selection differential, or from the proportion selected via selection intensity and the trait's standard deviation — plus generations to reach a target, gain per year with the generation interval, and the realised heritability from observed gain.

Selection specified as
Heritability h² (narrow sense)
Population mean
Phenotypic standard deviation
Proportion selected as parents
Selection differential S (differential mode)
Observed response R (realised mode)
Target mean to reach
Generation interval
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How it works

R = h² × S · S = i × σ_P (i = selection intensity from the proportion kept: 50% → 0.80, 20% → 1.40, 10% → 1.75, 5% → 2.06, 1% → 2.67) · gain per year = R ÷ generation interval · realised h² = R ÷ S

When breeders keep only the best animals or plants as parents, the next generation moves toward them, but only by the share of the parents' superiority that is genetic rather than environmental — that share is the narrow-sense heritability, and the breeder's equation multiplies it by the selection differential to give the response. How far above average the chosen parents are depends on how few are kept: truncating a normal distribution at the top 20% gives parents 1.4 standard deviations above the mean, the top 1% about 2.7, with steeply diminishing returns that also shrink the breeding population and raise inbreeding. Dividing by the generation interval converts response to annual gain, which is why dairy and poultry programmes chase shorter generations as hard as higher intensity.

Worked example

A trait with mean 100, standard deviation 10 and heritability 0.3: keeping the top 20% as parents gives an intensity of 1.40, a selection differential of 14 and a response of 0.3 × 14 = 4.2 per generation — 1.68 per year with a 2.5-year interval. Reaching a mean of 120 would take about 4.8 generations (12 years). If the observed response had been only 2.8, the realised heritability would be 0.2.

Frequently asked questions

Why is the response smaller than the differential?

Because most of what makes a parent outstanding is environment, dominance and luck that are not passed on. Only additive genetic differences are transmitted, and h² is exactly that fraction of the phenotypic variance.

Can I select on several traits?

Yes, but the total intensity is shared: selecting independently on n traits at the same overall proportion gives each roughly 1/√n of the single-trait response. Index selection weights traits by economic value and heritability to do better.

Related calculators

Textbook ecology, forestry and quantitative-genetics formulas with simplifying assumptions. Field constants (z, k, h², efficiencies) vary by system — use locally calibrated values for management or research decisions.